Let me throw out a question about the "duration" part.Dose = concentration X duration of exposure X frequency of exposure.
Some of the reading I've done points out the unique ability of velamen to "instantly" bond with mineral ions and hold onto them, presumably an evolutionary development to take advantage of the fact that the very first part of a rainstorm will contain the most nutrients, and to prevent the deluge that follows from washing it all away.
However, that says absolutely nothing about the rate of uptake, and that is what gives relative weight to the duration of exposure. If I run my fertilizer solution over a plant's roots for two minutes, is the plant getting twice the dose as it would at one minute? I doubt it.
Any nutrient uptake dynamics studies on orchids out there to lend us a hand?